Caroline Cappello

PhD Student



I graduated from Brandeis University with a bachelor’s degree in environmental studies and theater arts (though haven’t yet found a way to combine the two!). My focus on wildlife ecology and conservation as an undergraduate provided the opportunity to study large mammal ecology in Tanzania and Kenya. After graduating and before landing in the Boersma Lab, I wandered the country as a seasonal field biologist, working with a variety of avian and mammalian species. Some highlights from those nomadic years include monitoring elk behavior in Oregon, mist-netting yellow-billed cuckoos in the Sonoran Desert, tracking American martens through snowy Wisconsin forests, and managing breeding piping plover and least tern populations along the coast of southern Maine.

Scientific Interests

I am interested in the ability of species to persist in a changing environment. Specifically, I am interested in the factors that drive the timing of key life events like reproduction, the capacity of individuals to shift their reproductive phenology in response to changes in their environment, and the implications of these phenological shifts for the population as a whole. In the Boersma lab, I consider this topic in two systems:

Phenological shifts in seabird reproduction – Punta Tombo, Argentina–

Breeding at Punta Tombo, the world’s largest Magellanic penguin colony, is later and less synchronous than in the early 1980s. By combining the Boersma Lab’s 35-year ecological dataset with oceanographic and fisheries data, I am exploring the implications of these phenological shifts on reproductive success and breeding-adult recruitment to the colony.

Breeding biology and conservation of an endangered seabird – Galápagos Islands, Ecuador–

The Galápagos penguin is a rare seabird that, unlike many other penguin species, does not nest colonially. Locating these birds and understanding their natural history and population dynamics is a challenge. Along with Dee Boersma and Godfrey Merlen, I aim to understand the drivers of reproduction (e.g. nest-site availability, oceanographic conditions) for the Galápagos penguin and to provide information to Galápagos National Park that will aid them in the management of this small, endangered population.




Cappello, C. D. and P. D. Boersma. 2018. Sexing Galápagos penguins (Spheniscus mendiculus) by morphological measurements. Endangered Species Research. 35: 169-173

Boersma, P. D., C. D. Cappello, and G. Merlen. 2017. First observations of post-fledging care in Galápagos penguins (Spheniscus mendiculus). Wilson Journal of Ornithology. 129(1): 186-191

McNeil, S. E., D. Tracy, and C. D. Cappello. 2015. Loop migration and Chaco wintering by a Western Yellow-billed Cuckoo. Western Birds 46(3): 244-255